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Iwatsubo, Y. & N. Naruhashi. 1991. Karyomorphological and cytogenetical studies of Rubus parvifolius, R. coreanus and R. @xhiraseanus (Rosaceae). Cytologia 56: 151–156.
Pool, P. A., R. Ingram & R. J. Abbott. 1981. Karyotype variation in Rubus with particular reference to R. idaeus L. and R. coreanus Miquel. Cytologia 46: 125–132.